Sacalia

Genus of turtles

Sacalia

Sacalia quadriocellata
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Order:	Testudines
Suborder:	Cryptodira
Superfamily:	Testudinoidea
Family:	Geoemydidae
Subfamily:	Geoemydinae
Genus:	Sacalia Gray, 1870^[1]

Sacalia is a genus of turtles in the family Geoemydidae (formerly called Bataguridae). The genus was erected by John Edward Gray in 1870. The Sacalia is native to Southereastern China, Laos, and Vietnam. It was declared an endangered species of turtle by the IUCN Red List due to its illegal trade.^[2] The sacalia has a carapace of dark-brown color that permits it to camouflage to its habitat. They are believed to select the area of a river that better matches their color.^[3]

Species

Two species are recognized.^[1]

Sacalia bealei, Beale's eyed turtle or Beale's four-eyed turtle
Sacalia quadriocellata, four-eyed turtle

Hybridization

What was described as a new species, Sacalia pseudocellata, Chinese false-eyed turtle, is actually a hybrid of a male Cuora trifasciata and a female Sacalia quadriocellata.^[4]

References

Wikispecies has information related to Sacalia.

^ ^a ^b ITIS (Integrated Taxonomic Information System). www.itis.gov.
^ Le, M., McCormack, T., Hoang, H., Duong, H., Nguyen, T., Ziegler, T., Nguyen, H., & Ngo, H. (2020). Threats from wildlife trade: The importance of genetic data in safeguarding the endangered Four-eyed Turtle (Sacalia quadriocellata). Nature Conservation, 41(2), 91–111. https://doi.org/10.3897/natureconservation.41.54661
^ Xiao, F., Yang, C., Shi, H., Wang, J., Sun, L., & Lin, L. (2016). Background matching and camouflage efficiency predict population density in four-eyed turtle (Sacalia quadriocellata). Behavioural Processes, 131, 40–46. https://doi.org/10.1016/j.beproc.2016.08.007
^ Stuart, Bryan L. & Parham, James F. (2007). "Recent hybrid origin of three rare Chinese turtles". Conservation Genetics. 8 (1): 169-175.

Cheloniidae	†Allopleuron Caretta †Carolinochelys Chelonia †Eochelone Eretmochelys †Gigantatypus †Glarichelys †Itilochelys Lepidochelys †Mexichelys †Miocaretta Natator †Pacifichelys †Syllomus †Tasbacka
Dermochelyidae	†Arabemys †Corsochelys †Cosmochelys Dermochelys †Eosphargis †Mesodermochelys †Psephophorus
	†Euclastes †Peritresius †Procolpochelys †Protosphargis †Puppigerus

Kinosternoidea

Dermatemydidae	Dermatemys
Kinosternidae	Claudius †Hoplochelys Kinosternon Staurotypus Sternotherus

Testudinoidea

Emydidae	†Acherontemys Chrysemys Clemmys Deirochelys Emys Actinemys Emydoidea Glyptemys Graptemys Malaclemys Pseudemys Terrapene Trachemys †Wilburemys
Geoemydidae	Batagur †Banhxeochelys Cuora Cyclemys Geoclemys Geoemyda Hardella Heosemys Leucocephalon Malayemys Mauremys Melanochelys Morenia Notochelys Orlitia Pangshura Rhinoclemmys Sacalia Siebenrockiella Vijayachelys
Platysternidae	Platysternon
Testudinidae	Aldabrachelys Astrochelys Centrochelys Chelonoidis Chersina Cylindraspis †Cymatholcus †Floridemys Geochelone Gopherus †Hadrianus †Hesperotestudo Homopus Indotestudo Kinixys Malacochersus Manouria †Megalochelys †Oligopherus Psammobates Pyxis †Solitudo Stigmochelys †Stylemys Testudo

Trionychia

Carettochelyidae	†Allaeochelys †Anosteira Carettochelys
Trionychidae	Amyda Apalone †Axestemys Chitra Cyclanorbis Cycloderma †Drazinderetes Dogania †Gilmoremys †Hutchemys †Khunnuchelys Lissemys Nilssonia Palea †Palaeoamyda Pelochelys Pelodiscus Rafetus Trionyx
	†Basilochelys †Sinaspideretes

Chelydridae	Chelydra †Chelydrops †Chelydropsis †Emarginachelys †Macrocephalochelys Macrochelys †Planiplastron †Protochelydra
†Nanhsiungchelyidae	†Anomalochelys †Basilemys †Jiangxichelys
†Protostegidae	†Alienochelys †Archelon †Atlantochelys †Bouliachelys †Calcarichelys †Cratochelone †Desmatochelys †Iserosaurus †Notochelone †Ocepechelon †Pneumatoarthrus †Protostega †Rhinochelys †Santanachelys †Terlinguachelys
	†Adocus †Argillochelys †Bashuchelys †Ctenochelys †Prionochelys †Toxochelys

Pleurodira

†Araripemydidae	†Araripemys
†Bothremydidae	†Araiochelys †Arenila †Azabbaremys †Bothremys †Cearachelys †Chedighaii †Chupacabrachelys †Eotaphrosphys †Foxemys †Galianemys †Ilatardia †Inaechelys †Itapecuruemys †Jainemys †Kinkonychelys †Kurmademys †Labrostochelys †Nigeremys †Phosphatochelys †Polysternon †Puentemys †Rosasia †Rhothonemys †Sankuchemys †Taphrosphys †Ummulisani †Zolhafah
Chelidae	Acanthochelys Chelodina Chelus Elseya Elusor Emydura Hydromedusa †Lomalatachelys Mesoclemmys Myuchelys Phrynops Platemys †Prochelidella Pseudemydura Rheodytes Rhinemys †Yaminuechelys
Pelomedusidae	Pelomedusa Pelusios
Podocnemididae	†Albertwoodemys †Bauruemys †Brontochelys †Caninemys †Carbonemys †Cerrejonemys †Cordichelys Erymnochelys †Lapparentemys †Latentemys Peltocephalus Podocnemis †Stupendemys
†Sahonachelyidae	†Sahonachelys †Sokatra

	†Caribemys †Caririemys †Tacuarembemys

Phylogenetic arrangement of turtles based on Turtles of the World 2017 Update: Annotated Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status. † = extinct.

See also List of Testudines families

Taxon identifiers
Sacalia	Wikidata: Q2396266 Wikispecies: Sacalia ADW: Sacalia BOLD: 191065 CoL: 63RL7 EoL: 35997 GBIF: 2443456 iNaturalist: 39920 IRMNG: 1418004 ITIS: 551759 NCBI: 74933 Open Tree of Life: 105475 Paleobiology Database: 137545